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Genus Enicospilus. Enicospilus purgatus? Class: Insecta Subclass: Pterygota Order: Hymenoptera Suborder: Apocrita Superfamily: Ichneumonoidea Family: Ichneumonidae (Ichneumonid Wasps) Subfamily: Ophioninae (Short-tailed Ichneumonid Wasps) Tribe: Enicospilini Genus: Enicospilus Color & Size: It has a distinctive slender, orange to yellowish-brown body and can be quite large, often around 1 inch long. Eyes & Antennae: Note the very large, dark compound eyes and exceptionally long, multi-segmented antennae, which are adaptations for its nocturnal lifestyle. Wings: It has two pairs of wings, which distinguishes it from a similar-looking crane fly, a type of true fly with only one pair. "Horse Head" Wing Pattern: A defining characteristic of this family (Ichneumonidae) is a specific "horse-head" shape formed by the veins in the forewings. Enicospilus is a genus of large Ichneumonidae wasp, which do not sting. They are unusual for wasps in that they fly at night and can be found near porch lights. While these wasps resemble crane flies with long legs and long, thin bodies, the presence of two sets of wings distinguishes them from the single pair of wings of diptera (true flies including crane flies). There are almost 700 described species of Enicospilus.... nocturnal parasitoid wasps within the family Ichneumonidae and subfamily Ophioninae. Parasitic Nature: Enicospilus are koinobiont endoparasitoids, primarily attacking the larvae of Lepidoptera (moths and butterflies). Nocturnal Activity: Unlike many other wasps, Enicospilus are crepuscular or nocturnal, frequently attracted to light traps during field surveys. Body: Large, slender, and typically yellowish-brown (testaceous) in color. Eyes & Ocelli: Markedly large eyes and ocelli, adapted for nocturnal vision. Wings: Presence of a large fenestra (a clear area) in the fore wing discosubmarginal cell, often containing one or more hardened, pigmented sclerites. Mandibles: Strongly narrowed and twisted apically. functioning as a natural regulator of agricultural and forest pests through its specialized parasitic behavior. 1. Specialized Parasitic Behavior Koinobiont Strategy: Enicospilus wasps are koinobiont endoparasitoids. The female uses its short ovipositor to inject a single egg into a host caterpillar. Unlike idiobionts that paralyze their host immediately, koinobionts allow the caterpillar to continue feeding and growing, providing a living environment for the developing wasp larva. Host Specificity: These wasps primarily target large larvae of Lepidoptera (moths and butterflies). Research indicates that in tropical regions like India, Enicospilus species often specialize on a single host species to overcome complex chemical defenses in tropical plants. Nocturnal Activity: Highly unusual for wasps, Enicospilus are nocturnal or crepuscular. Their large eyes and ocelli are adapted for hunting host larvae at night when many moth species are most active. Natural Pest Control: By parasitizing moth larvae, Enicospilus wasps act as vital biological control agents. They regulate populations of pests that would otherwise damage crops and forests. Harmless to Humans: Despite their large size (often reaching 1 inch), they are considered stingless or harmless to humans because they do not have a colony-defending instinct like social wasps. Harmless to Humans: Despite their intimidating appearance and size, they are stingless to humans. The female possesses an ovipositor (egg-laying tube) rather than a stinger, so they do not defend a colony or attack people. Natural Pest Controller: They are beneficial parasitic wasps, acting as natural biological control agents by laying their eggs inside the larvae (caterpillars) of various large moths. Life Cycle Summary Egg: Laid inside the host caterpillar. Larva: Feeds internally on the host's hemolymph, carefully avoiding vital organs to keep the host alive for as long as possible. Pupa: Once the larva is mature, it emerges from the host (killing it) and spins a distinctive, often fibrous cocoon nearby. Adult: Emerges to feed on nectar, honeydew, and water, focusing primarily on reproduction and finding new hosts. frequently attracted to lights, which is likely how I spotted it. #Enicospilus #wasp #nocturnalwasp #koinobiontendoparasitoids #endoparasitoids #Ichneumon #Ichneumonidae #parasitoid