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Spring Mechanisms in the Foot and Lower Extremity In this 18-minute mini-lecture, I review how the spring mechanisms within the foot and lower extremity are vital to optimizing the biomechanics of weightbearing activities. Hooke's Law is reviewed along with the concepts of spring stiffness, viscoelasticity, and how running, galloping, and hopping animals all use elastic strain energy stored within their muscle, tendons and ligaments in order to improve the metabolic efficiency, speed, power and safety of their locomotor activities. In addition, the spring-like nature of the midtarsal joint, midfoot joints and metatarsal rays are explored, including the important concept that a "locking midtarsal joint" is neither a reality nor a desirable biomechanical feature of the human foot. Rather, the spring-like nature of the midtarsal joint, midfoot joints and metatarsal rays are all biomechanically necessary to allow the bipedal human to help maintain their metatarsal heads in a plantigrade position on a wide variety of uneven surfaces in order to improve stability of the foot, and rest of the body, during the multitude of weightbearing activities that we perform on a daily basis.