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Please enjoy this video examining the early evolution of Pterosaurs, the first vertebrate animals to develop powered flight. Thumbnail art by Mark Witton. / drpolaris https://www.deviantart.com/drpolaris Sources used for the video: Baron, Matthew G (2020). "Testing pterosaur ingroup relationships through broader sampling of avemetatarsalian taxa and characters and a range of phylogenetic analysis techniques". PeerJ. 8: e9604. Bestwick, Jordan; Unwin, David M.; Butler, Richard J.; Purnell, Mark A. (28 October 2020). "Dietary diversity and evolution of the earliest flying vertebrates revealed by dental microwear texture analysis". Nature Communications. 11 (1): 5293. Claessens LP, O'Connor PM, Unwin DM (2009). Sereno P (ed.). "Respiratory evolution facilitated the origin of pterosaur flight and aerial gigantism". PLOS ONE. 4 (2): e4497. Codorniú, Laura; Paulina Carabajal, Ariana; Pol, Diego; Unwin, David; Rauhut, Oliver W.M. (2016). "A Jurassic pterosaur from Patagonia and the origin of the pterodactyloid neurocranium". PeerJ. 4: e2311. Dyke GJ, Nudds RL, Rayner JM (July 2006). "Limb disparity and wing shape in pterosaurs". J. Evol. Biol. 19 (4): 1339–42. Elgin RA, Hone DW, Frey E (2011). "The Extent of the Pterosaur Flight Membrane". Acta Palaeontologica Polonica. 56 (1): 99–111. Garcia, Maurício S.; Müller, Rodrigo T. (2025). "Triassic pterosaur precursors of Brazil: catalog, evolutionary context, and a new hypothesis for phylogenetic relationships of Pterosauromorpha". Anais da Academia Brasileira de Ciências. 97 (suppl 1). Müller, R. T.; Ezcurra, M. D.; Garcia, M. S.; Agnolín, F. L.; Stocker, M. R.; Novas, F. E.; Soares, M. B.; Kellner, A. W. A.; Nesbitt, S. J. (2023). "New reptile shows dinosaurs and pterosaurs evolved among diverse precursors". Nature. 620 (7974): 589–594. Pittman, Michael; Barlow, Luke A.; Kaye, Thomas G.; Habib, Michael B. (2021). "Pterosaurs evolved a muscular wing–body junction providing multifaceted flight performance benefits: Advanced aerodynamic smoothing, sophisticated wing root control, and wing force generation". Proceedings of the National Academy of Sciences. 118 (44): e2107631118. Prondvai, E.; Stein, K.; Ősi, A.; Sander, M. P. (2012). Soares, Daphne (ed.). "Life history of Rhamphorhynchus inferred from bone histology and the diversity of pterosaurian growth strategies". PLOS ONE. 7 (2): e31392. Unwin, David M.; Martill, David M. (December 2020). "No protofeathers on pterosaurs". Nature Ecology & Evolution. 4 (12): 1590–1591. Vecchia, Dalla; Marco, Fabio (2019). "Seazzadactylus venieri gen. et sp. nov., a new pterosaur (Diapsida: Pterosauria) from the Upper Triassic (Norian) of northeastern Italy". PeerJ. 7: e7363. Witmer LM, Chatterjee S, Franzosa J, Rowe T (2003). "Neuroanatomy of flying reptiles and implications for flight, posture and behaviour" (PDF). Nature. 425 (6961): 950–53. Witton, Mark P. (2013). Pterosaurs: Natural History, Evolution, Anatomy. Princeton University Press. Witton, Mark P. (2015). "Were early pterosaurs inept terrestrial locomotors?". PeerJ. 3: e1018. Yang, Zixiao; Jiang, Baoyu; Benton, Michael J.; Xu, Xing; McNamara, Maria E.; Hone, David W. E. (2023-07-26). "Allometric wing growth links parental care to pterosaur giantism". Proceedings of the Royal Society B: Biological Sciences. 290 Yang, Zixiao; Jiang, Baoyu; McNamara, Maria E.; Kearns, Stuart L.; Pittman, Michael; Kaye, Thomas G.; Orr, Patrick J.; Xu, Xing; Benton, Michael J. (January 2019). "Pterosaur integumentary structures with complex feather-like branching". Nature Ecology & Evolution. 3 (1): 24–30.